Calanus finmarchicus is a tiny marine crustacean that plays an outsized role in the North Atlantic Ocean.
It is a species of copepod—a small animal that forms part of the zooplankton drifting and swimming through the ocean.
An adult Calanus finmarchicus is generally only about 2 to 3 millimeters long, yet enormous populations of these animals help support some of the North Atlantic's most important marine ecosystems.1, 2
They feed largely on microscopic marine plants and, in turn, become food for fish and other animals. In effect, Calanus finmarchicus helps transfer energy produced by phytoplankton into the animals higher in the marine food web.
It also possesses an unusual adaptation for surviving the dramatic seasonal changes of northern waters: it stores large quantities of energy as wax esters.
Those wax ester-rich energy reserves are also what make Calanus finmarchicus the natural source of the marine oil known as Calanus oil.
Calanus finmarchicus: The Quick Answer
Scientific name: Calanus finmarchicus
Animal type: Copepod, a small marine crustacean
Adult size: Approximately 2–3 millimeters
Primary habitat: North Atlantic Ocean, including the Norwegian Sea
Primary food: Phytoplankton and other microscopic particles
Ecological role: Transfers energy from microscopic primary producers to fish and other marine animals
Major energy reserve: Wax esters
Notable fatty acids: EPA, DHA and SDA, among many others
Commercial significance: Natural source of Calanus oil
What Is a Copepod?
Copepods are small crustaceans found in marine and freshwater environments throughout the world.
That means Calanus finmarchicus belongs to the same broad animal group as familiar crustaceans such as shrimp, crabs and lobsters, although copepods are vastly smaller.
Calanus finmarchicus belongs to the genus Calanus and the family Calanidae.
It is part of the zooplankton, the diverse community of small animals living in the water column.
The word “plankton” sometimes gives the impression that these organisms simply float passively. In reality, copepods can swim and perform substantial vertical migrations through the water column, although their large-scale geographic movement is strongly influenced by currents.
What makes Calanus finmarchicus especially significant is not its individual size, but its enormous abundance and its central position in the North Atlantic food web.
Where Does Calanus finmarchicus Live?
Calanus finmarchicus is fundamentally a North Atlantic species.
A major scientific review of its distribution documented populations across much of the North Atlantic, from waters near the Gulf of Maine and eastern Canada across the Atlantic to Iceland, the waters around the Faroe Islands, the North Sea and the Norwegian Sea.1
Important population and overwintering centers have been identified in areas including:
- the Labrador Sea
- the Irminger Basin
- the Iceland Basin
- the Faroe-Shetland Channel
- the Norwegian Sea
- the Norwegian Trench
- deep basins of the Gulf of Maine
Ocean currents play an important part in its distribution. Populations can be transported considerable distances as water masses circulate through the North Atlantic.
The southern Norwegian Sea is particularly important because deep waters there provide a major center for production and overwintering populations. 1
What Does Calanus finmarchicus Look Like?
To the naked eye, an individual Calanus finmarchicus is tiny. Adults typically measure only about 2–3 millimeters in length. 2
Under magnification, it is clearly recognizable as a small crustacean. It has a translucent, segmented body, long antennae and multiple appendages used in swimming and feeding.
One particularly striking feature can become visible in later developmental stages:
a large internal oil sac.
This oil sac may occupy a substantial portion of the animal's body cavity when its energy reserves are high.
Unlike the fat storage familiar in many terrestrial animals, much of the stored lipid in late-stage Calanus finmarchicus consists of wax esters.
The Life Cycle of Calanus finmarchicus
The life cycle of Calanus finmarchicus is closely synchronized with the seasonal productivity of the North Atlantic.
After hatching from an egg, the animal progresses through six naupliar stages.
It then develops through five juvenile copepodite stages, commonly labeled CI through CV, before reaching the sixth copepodite stage—the adult male or female.1, 2
The first two naupliar stages do not feed. As development continues, the young copepods begin feeding and growing.
Within much of its main North Atlantic range, the life cycle is approximately annual, although timing and number of generations can vary with geography, temperature and food availability.1
This life cycle contains an extraordinary seasonal strategy.
During the productive part of the year, young Calanus feed, grow and accumulate energy. Later-stage animals can then descend hundreds of meters into deeper water and spend an extended period in a low-metabolism state before returning toward the surface as another productive season begins.
Following the Spring Phytoplankton Bloom
To understand Calanus finmarchicus, it helps to understand the North Atlantic spring bloom.
During winter at high northern latitudes, limited sunlight restricts the growth of phytoplankton.
As daylight increases in spring and conditions become favorable, microscopic phytoplankton can reproduce rapidly, producing a major pulse of biological productivity.
Calanus is adapted to take advantage of this seasonal abundance.
Research has shown a close relationship between Calanus development and the timing of phytoplankton production. 1, 3
By consuming phytoplankton, Calanus takes energy originally captured from sunlight and incorporates it into its own tissues and lipid reserves.
Sunlight → Phytoplankton → Calanus finmarchicus → Fish and other marine animals
This is one reason Calanus occupies such an important position in northern marine ecosystems.
Why Does Calanus finmarchicus Store Wax Esters?
One of the defining biological characteristics of Calanus finmarchicus is its ability to accumulate large amounts of lipid as wax esters.
A wax ester is formed when a fatty acid is chemically linked to a long-chain fatty alcohol.
This differs from a triglyceride, in which three fatty acids are attached to glycerol.
Studies of Calanus finmarchicus have shown that wax esters become especially important as the animal matures and prepares for its extended period at depth. 3, 4, 5
Wax esters provide a highly concentrated long-term energy reserve.
Researchers studying Arctic and sub-Arctic Calanus species have described them as important converters of energy from phytoplankton into energy-rich wax esters that can then move through the marine food web. 3
The Remarkable Calanus Oil Sac
As later-stage Calanus finmarchicus feed during the productive season, they accumulate lipid inside a conspicuous internal oil sac.
Lipid accumulation begins during the copepodite stages and becomes especially substantial in the later stages as the animal prepares for overwintering. 2
In lipid-rich individuals, this oil sac can occupy a large portion of the body cavity.
Wax esters may account for the great majority of stored lipid in these late-stage animals. Research has reported wax-ester proportions reaching close to 90% of total lipid under some conditions. 5
The amount and composition are not constant throughout the year. They vary with developmental stage, season, food supply and geographic location.
This natural oil-storage system is ultimately the biological origin of the wax ester-rich composition found in commercial Calanus oil.
How Calanus Survives the North Atlantic Winter
One of the most remarkable parts of the Calanus life cycle occurs far below the ocean surface.
Many individuals in the fifth copepodite stage, known as CV, descend into deeper water and enter a period of greatly reduced activity and metabolism.
Scientists commonly refer to this dormant or resting period as diapause.
Depending on region, overwintering Calanus may occur hundreds of meters below the surface and, in deep ocean basins, at depths exceeding 1,000 meters.1, 6
During this period, feeding is greatly reduced or absent and metabolism slows substantially.
The animal survives largely on the lipid reserves accumulated during the productive months above.
Those reserves are predominantly wax esters.
When the seasonal cycle turns again, surviving animals return toward productive surface waters, complete development and reproduce.
The wax ester is therefore not an incidental part of Calanus biology. It is closely connected to the species' survival strategy.
Why Is Calanus finmarchicus So Important to the Marine Food Web?
Scientists frequently describe Calanus finmarchicus as a key species in the North Atlantic pelagic ecosystem. 1, 7
The reason is its position between microscopic primary producers and larger animals.
Phytoplankton capture energy through photosynthesis. Calanus consumes phytoplankton and packages some of that energy into nutrient-rich animal biomass and lipids.
Calanus then becomes prey for numerous animals.
In the Norwegian Sea, it is an important food source for commercially important pelagic fish including:
- herring
- mackerel
- blue whiting
Calanus and related zooplankton are also important prey for fish larvae, juvenile fish and other marine predators.
The Norwegian Institute of Marine Research describes Calanus finmarchicus as a keystone species in the Norwegian Sea ecosystem.7
In other words, a tiny animal only a few millimeters long helps support organisms millions of times larger than itself.
How Abundant Is Calanus finmarchicus?
Calanus finmarchicus is one of the dominant mesozooplankton species of the North Atlantic and is particularly abundant in the Norwegian Sea.1, 8
Measuring an animal that is only a few millimeters long across an ocean basin is difficult, so estimates vary according to methodology, region and year.
Nevertheless, scientific modeling demonstrates the extraordinary scale of the resource.
A Norwegian Sea ecosystem model published in 2012 estimated annual Calanus finmarchicus production at approximately 29 million metric tons of carbon. 8
Norwegian resource assessments and management materials have also cited estimates on the order of approximately 190–290 million metric tons of wet-weight annual production in the Norwegian Sea.
These figures describe production across an immense ecosystem; they should not be confused with the amount commercially harvested.
Commercial catches represent only a fraction of the estimated natural production and are subject to Norwegian management rules and quotas.
Where Do Its Omega-3 Fatty Acids Come From?
The story begins lower in the food web.
Marine primary producers such as phytoplankton are fundamental sources of fatty acids that move through ocean ecosystems.
As Calanus feeds, it acquires and metabolically modifies dietary lipids and stores a substantial portion of its energy in wax esters.
Scientific analyses of Calanus finmarchicus and its oil have identified numerous fatty acids, including the omega-3 fatty acids:
- EPA — eicosapentaenoic acid
- DHA — docosahexaenoic acid
- SDA — stearidonic acid
Unlike conventional fish oil, much of the fatty-acid content of Calanus is stored within wax ester molecules rather than triglycerides.
The long-chain fatty alcohol portion of the wax esters is also an important part of this unusual lipid system.
This is why Calanus oil differs chemically from conventional fish and krill oils even though all three can provide marine omega-3 fatty acids.
How Does Calanus finmarchicus Become Calanus Oil?
Calanus oil is the lipid extract obtained from Calanus finmarchicus.
Because the animal naturally stores so much of its lipid as wax esters, the resulting oil is also predominantly wax ester-based.
Published scientific reviews of commercial Calanus oil describe approximately 80–90% of its lipid fraction as wax esters, although natural composition varies. 9
These wax esters contain both fatty acids and long-chain fatty alcohols.
The oil also naturally contains a broad spectrum of fatty acids and carotenoid compounds including astaxanthin.
Calanus oil is therefore not simply purified EPA and DHA extracted from another marine animal.
It reflects the distinctive lipid-storage biology of Calanus finmarchicus.
How Does Zooca Harvest Calanus finmarchicus?
The Calanus oil used in Arctic Ruby Oil is supplied by Zooca® / Calanus AS of Norway.
Zooca has spent more than two decades developing technology for harvesting and processing Calanus finmarchicus.
According to Zooca, the company developed a patented trawl specifically for this unusually small marine resource. 10
Rather than using conventional fishing gear designed for fish, the harvesting system uses a wide, shallow trawl with mesh designed for collecting Calanus while allowing water to pass through.
Zooca reports that its commercial harvesting season generally runs from approximately May through mid-August in Norwegian waters, although locations and timing vary because Calanus distribution changes with oceanographic conditions.
The harvested material is frozen aboard the vessel before processing.
Zooca completed a dedicated Calanus finmarchicus processing facility in Sortland, Northern Norway, in 2021.
What About Sustainability?
The abundance of Calanus finmarchicus makes it an intriguing marine resource, but abundance alone does not make unrestricted harvesting appropriate.
Calanus is ecologically important precisely because so many other animals depend upon it.
For that reason, commercial harvesting needs to account for both the size of the resource and its function within the food web.
Norway moved from experimental harvesting to a regulated commercial Calanus finmarchicus fishery beginning in 2019.
The fishery operates within Norway's marine-resource management framework, which includes permits, quotas and precautionary management principles.
Zooca states that its harvesting operations comply with these allocated quotas and that its specialized trawling technology is intended to minimize bycatch.10
Scientific evaluation remains important. Earlier reviews of Calanus utilization specifically emphasized that ecological sustainability should be assessed carefully before large-scale expansion of the fishery. 2
That precaution is appropriate for a species that plays such an important role in the North Atlantic ecosystem.
Calanus finmarchicus and a Changing North Atlantic
Calanus finmarchicus is also important to scientists studying climate-driven changes in the North Atlantic.
Its development, distribution and reproductive success are influenced by factors including:
- water temperature
- ocean circulation
- phytoplankton abundance
- timing of the spring bloom
- seasonal light conditions
Changes in temperature can alter where different Calanus species thrive.
In parts of the northeastern Atlantic and North Sea, warming has been associated with changes in the relative distribution of C. finmarchicus and the more warm-water-associated Calanus helgolandicus.
Because Calanus is a major food source for many fish, changes in its abundance or seasonal timing can potentially affect organisms higher in the food web.
This is one reason Calanus finmarchicus has been studied so extensively by oceanographers and marine ecologists.
What Does Calanus finmarchicus Have to Do With Arctic Ruby Oil?
Calanus finmarchicus is the marine organism from which the Calanus oil in Arctic Ruby Oil is derived.
That distinction matters because Arctic Ruby Oil is therefore not conventional fish oil and it is not krill oil.
Its underlying marine lipid comes from a different organism with a different energy-storage system.
Calanus survives the extreme seasonality of the North Atlantic in part by accumulating large stores of wax ester-rich lipid.
Those naturally occurring lipids contain:
- EPA
- DHA
- SDA
- other marine fatty acids
- long-chain fatty alcohols
- naturally occurring astaxanthin
Zooca® extracts this natural lipid from Calanus finmarchicus for use as Calanus oil.
Arctic Ruby Oil uses that Zooca® Calanus® Oil as its defining marine oil ingredient.
Calanus finmarchicus → wax ester-rich natural lipid → Zooca® Calanus® Oil → Arctic Ruby Oil
Frequently Asked Questions
Is Calanus finmarchicus a fish?
No. Calanus finmarchicus is a copepod, which is a tiny crustacean. Although its oil provides marine omega-3 fatty acids, the organism itself is not a fish.
Is Calanus finmarchicus krill?
No. Calanus and krill are both small marine crustaceans and both are part of marine plankton communities, but they are different animals. Calanus oil is particularly notable for its wax ester-rich lipid structure, while krill oil contains a substantial phospholipid fraction.
How big is Calanus finmarchicus?
Adult Calanus finmarchicus are generally around 2–3 millimeters long. Despite their tiny size, collectively they represent an enormous amount of biological production in the North Atlantic.
Where is Calanus finmarchicus found?
It occurs across much of the North Atlantic, with major populations in regions including the Labrador Sea, Icelandic waters, the northeastern Atlantic and the Norwegian Sea.
What does Calanus finmarchicus eat?
Calanus feeds extensively on phytoplankton, including diatoms and other microscopic primary producers, although its diet can vary with environmental conditions and life stage.
What animals eat Calanus finmarchicus?
Calanus is an important prey species for many fish and other marine animals. In the Norwegian Sea it is an especially important food source for species such as herring, mackerel and blue whiting, and it also supports fish larvae and juvenile fish.
Why does Calanus finmarchicus contain so much oil?
Calanus accumulates lipid during productive periods so that it can survive long periods when food is scarce. Much of that stored energy takes the form of wax esters contained in a prominent internal oil sac.
Why are wax esters important to Calanus finmarchicus?
Wax esters function as long-term energy reserves. They help support Calanus during its seasonal migration into deep water and its extended period of reduced activity and feeding.
Does Calanus finmarchicus naturally contain omega-3 fatty acids?
Yes. Its lipid contains numerous fatty acids, including the omega-3 fatty acids EPA, DHA and SDA. Much of this fatty-acid content is naturally incorporated into wax ester molecules.
Is Calanus oil the same as fish oil?
No. Fish oil is obtained from fish and is generally dominated by triglyceride-based lipids in its natural form. Calanus oil comes from the copepod Calanus finmarchicus and is predominantly wax ester-based.
What is the difference between Calanus finmarchicus and Calanus oil?
Calanus finmarchicus is the living marine organism. Calanus oil is the lipid extracted from that organism.
What is the relationship between Calanus finmarchicus and Arctic Ruby Oil?
Arctic Ruby Oil contains Zooca® Calanus® Oil derived from Calanus finmarchicus. The species' natural wax ester-rich lipid system is therefore the biological origin of the distinctive marine oil used in Arctic Ruby Oil.
The Bottom Line
Calanus finmarchicus is a tiny North Atlantic copepod with an exceptionally large ecological role.
It feeds on microscopic primary producers and, in turn, becomes a major food source for fish and other marine animals.
Its life is closely synchronized with the dramatic seasonal cycle of northern waters.
During productive periods, Calanus accumulates a large reserve of energy-rich lipids. Much of that energy is stored as wax esters inside a prominent oil sac.
These reserves help the animal survive an extended period at depth when feeding and metabolism are greatly reduced.
The same unusual lipid-storage system gives Calanus oil its defining chemical characteristic:
a predominantly wax ester-based marine oil containing omega-3 fatty acids together with long-chain fatty alcohols and other naturally occurring marine lipids.
That is why understanding Calanus finmarchicus is essential to understanding what makes Calanus oil—and Arctic Ruby Oil—different from conventional fish and krill oils.
Scientific References
- Melle W, Runge JA, Head E, Plourde S, Castellani C, Licandro P, Pierson J, Jónasdóttir SH, Johnson C, Broms C, et al. “The North Atlantic Ocean as Habitat for Calanus finmarchicus: Environmental Factors and Life History Traits.” Progress in Oceanography. 2014;129:244–284. DOI: 10.1016/j.pocean.2014.04.026.
- Eysteinsson ST, Gudjónsdóttir M, Jónasdóttir SH, Arason S. “Review of the Composition and Current Utilization of Calanus finmarchicus – Possibilities for Human Consumption.” Trends in Food Science & Technology. 2018;79:10–18. DOI: 10.1016/j.tifs.2018.06.019.
- Falk-Petersen S, Mayzaud P, Kattner G, Sargent JR. “Lipids and Life Strategy of Arctic Calanus.” Marine Biology Research. 2009;5(1):18–39. DOI: 10.1080/17451000802512267.
- Fraser AJ, Sargent JR, Gamble JC, Seaton DD. “Formation and Transfer of Fatty Acids in an Enclosed Marine Food Chain Comprising Phytoplankton, Zooplankton and Herring.” Published research examining marine lipid transfer through planktonic food webs.
- Fraser AJ, Sargent JR, Gamble JC. “Lipid Class and Fatty Acid Composition of Calanus finmarchicus, Pseudocalanus sp. and Temora longicornis from a Nutrient-Enriched Seawater Enclosure.” Journal of Experimental Marine Biology and Ecology. 1989;130(1):81–92. DOI: 10.1016/0022-0981(89)90020-8.
- Head EJH, Pepin P. Research examining seasonal and depth-dependent lipid reserves and diapause strategies of Calanus finmarchicus populations in the Northwest Atlantic. See also the extensive diapause literature summarized by Melle et al. and Falk-Petersen et al.
- Institute of Marine Research, Norway. “Calanus” and “Norwegian Sea” species and ecosystem resources. The Institute identifies Calanus finmarchicus as a key component of the Norwegian Sea ecosystem and an important prey species for pelagic fish.
- Hjøllo SS, Huse G, Skogen MD, Melle W. “Modelling Secondary Production in the Norwegian Sea With a Fully Coupled Physical/Primary Production/Individual-Based Calanus finmarchicus Model System.” Marine Biology Research. 2012;8(5–6):508–526. DOI: 10.1080/17451000.2011.642805.
- Schots PC, Pedersen AM, Eilertsen KE, Olsen RL, Larsen TS. “Possible Health Effects of a Wax Ester Rich Marine Oil.” Frontiers in Pharmacology. 2020;11:961. DOI: 10.3389/fphar.2020.00961.
- Zooca® / Calanus AS, Norway. Current company information regarding Calanus finmarchicus harvesting, patented harvesting technology, processing, seasonality and regulatory compliance under Norway's marine resource management framework.